Cheers for the Photos.
I may have given you a bit of a bum steer a page or so back. I did conclude that the issues you're having were in the gearbox, largely based on the fact that you could engage reverse without grinding, but now I see this, I'd say the primary gear bushes are the issue - sorry for my bad.
I'm guessing here that when you've engaged reverse, the engine, oil and primary gear have been fairly cool / cold and this would likely be why there was no grinding for reverse, but as parts and oil have warmed up, the primary gear bush has just started picking up. I think you dodged a bullet here as from those I know of who have had trouble with these bushes (and that's been everyone who's fitted them) usually don't find out there's an issue until the gear is locked on the crank, by which time, the crank is karput (eaten by the bushes). These are simply too hard - your photos here shows that clear as day.
Since the gearbox is off, may as well check it over. I doubt you'd need to dis-assemble it any further to do this, maybe even get someone experienced who's local to check it too.
As for the Primary Gear Bushes, the factory found the answer here back in 1963-ish. They did try Bronze based bushes of various allying types but settled on a steel backed soft bush for one and a deva for the other. At least when these bushes go wrong (which isn't often), it's only the bushes that let go and the crank is nearly always saved. This is one of the primary design features of any bush. The Front Bush in a Genuine type is still available, they were (are) made by Glacier, you'll see their stamp on the back (G in a box). I have Deva bushes here, however I believe AC Dodd also has a small batch made too.