Center cap the right way around?
^ This was my first thought, my next is what condition are the hollow dowels in ?
There are a couple of nicks on them where they were gripped to pull them out, prior to machining the block.
They need to be perfectly clean.they are there to locate the cap absolute. the nicks can easily push it off. Even as much as 1/20th of one thousandth of an inch is enough to give issues. Assemble and Torque the caps to the block, the right way around, without the crank of bearing shells. While not the ultimate way of testing, feel with your finger or even the edge of a 0.001" feeler gauge across all the cap to block mating points to see if there any step there. Check from the cap to the block and the block to the cap. They must align firm on the dowels and have no step.